Abstract
The early Miocene sites of Moruorot and Kalodirr (Kenya, 17.5 Myr) have yielded a rich collection of mammals. New listriodont material from these localities, including a complete skull and a partial mandible, provide long awaited information on cranial features of early bunodont Listriodontinae. The evolution and systematics of the group are highly debated, especially regarding its first representatives. The new material described here sheds light on the differentiation of bunodont Listriodontinae in Africa and clarifies the systematics of the group. The first phylogenetic analysis of the Listriodontinae is here performed and supports close relationships between Kubano-choerus and a clade (Eurolistriodon, Listriodon). Lopholistriodon is the most basal representative of the listri-odontine clade. These first results stress the role of the African continent in the biogeographical history of the Listriodontinae.