Abstract
The lower Pleistocene Kritika Member of the Rhodes Formation is composed of predominantly siliciclastic deposits that are sometimes intercalated with hardbottoms formed by drowning of former beachrock beds. A wide variety of sclerobionts occur in this environment, very different from the poorest soft‐bottom populations occurring in other levels of the Kritika Member. The most characteristic, abundant and frequently diverse are the encrusting forms: coralline algae, bivalves, serpulid worms, and bryozoans. Numerous other groups of sessile and vagile organisms are associated with these crust‐forming communities. A number of skeletal organisms (sometimes including zooxanthellate coral colonies) formed small carbonate buildups on top of several beachrock beds. Boring organisms also occur. Evidence for a two‐step deepening succession over the previously formed beachrock is provided by palaeontological and sedimentological data. The rapidly cemented sandy‐conglomeratic beds were first colonised by a relatively shallow‐water (around 20 m) biocoenosis of mostly encrusting organisms. Another association, with encrusting and erect organisms, was later established in a deeper environment (20-40 m). A comparison with Holocene beachrock occurring on the coast of Rhodes was also undertaken. Their submerged part provided habitat for poorer but similar biotas to those occurring in the Kritika Member. The lower Pleistocene beachrock and associated organogenic fossil assemblages indicate repeated drowning episodes during deposition of the Kritika Member and provide help in sequence stratigraphic interpretations of these siliciclastic deposits. Notably, they can be used as indicators of relative sea‐level changes.