Abstract
The second Mesozoic representative of the (Dasyheleinae & Forcipomyiinae) clade of Ceratopogonidae, Devalquia brisaci gen. et sp. n., as well as the first Mesozoic Ceratopogoninae Heteromyiini Metahelea roggeroi sp. n., are the first arthropods described from the Santonian amber outcrop of Piolenc (Vaucluse, France). Both confirm the great morphological stability through time within the Ceratopogonidae. They also suggest that the other ceratopogonine clades were also present by at least the Upper Cretaceous.